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Clade comprising all crustaceans and hexapods
Pancrustacea is a major clade of arthropods that comprises all crustaceans and hexapods (insects and their entognath relatives). This grouping is contrary
Pancrustacea
Superclass of arthropods
'different, foreign' and καρίς/karis 'shrimp, crustacean') is a clade of Pancrustacea, containing Hexapoda (all insects, springtails & their close relatives)
Allotriocarida
Clade of arthropods
extant groups Myriapoda (millipedes and centipedes, among others) and Pancrustacea (including crustaceans and hexapods, the latter group containing insects)
Mandibulata
Group of arthropods
crustaceans, with the two groups now combined in the monophyletic clade Pancrustacea. The three classes Cephalocarida, Branchiopoda, and Remipedia are more
Crustacean
Phylum of invertebrates with jointed exoskeletons
that Crustacea and Hexapoda form a clade, Pancrustacea. The position of Myriapoda, Chelicerata and Pancrustacea remains unclear as of April 2012[update]
Arthropod
Subphylum of arthropods
of legs. Most recent studies have recovered Hexapoda as a subgroup of Pancrustacea as members of Allotriocarida. Hexapods have bodies ranging in length
Hexapoda
Class of crustaceans
; Koenemann, Stefan (2022). "Ostracoda". Evolution and Phylogeny of Pancrustacea. pp. 119–140. doi:10.1093/oso/9780195365764.003.0010. ISBN 978-0-19-536576-4
Ostracod
Geologic formation in France
The Argiles d'lignite du Soissonnais is a geologic formation in the Oise department of northern France. The formation has provided fossil mammals, reptiles
Argiles d'lignite du Soissonnais
Argiles_d'lignite_du_Soissonnais
Extinct order of crustaceans
doi:10.1086/274005. JSTOR 2450109. "A second enigmatic alsasuacaridid (Pancrustacea, Cyclida) from the Maastrichtian type area in The Netherlands (Late Cretaceous)"
Cyclida
Order of arthropods
hexapods (such as insects) if they are considered to fall under Crustacea (Pancrustacea). Terrestrial isopods play an important role in many tropical and temperate
Isopoda
Extinct Cambrian arthropod
predator. It is hesitantly presumed to be an early offshoot under the clade Pancrustacea in original description. Other organisms, including Henningsmoenicaris
Cambropachycope
Proposed group of arthropods
proposition holds true, the Myriochelata are the sister clade to the Pancrustacea (also Tetraconata), comprising classic crustaceans and hexapods. The
Myriochelata
Freshwater rift lake in Russia
R.; Koenemann, Stefan (2 November 2021). Evolution and Phylogeny of Pancrustacea: A Story of Scientific Method. Oxford University Press. ISBN 978-0-19-009598-7
Lake_Baikal
Class of arthropods
Chelicerata (sea spiders, horseshoe crabs, and arachnids) Mandibulata Myriapoda (centipedes, millipedes, and allies) Pancrustacea (crustaceans and hexapods)
Arachnid
Superclass of crustaceans
the phylogenetic affinities of the enigmatic thylacocephalans within Pancrustacea". Proceedings of the Royal Society B: Biological Sciences. 292. Collette
Multicrustacea
First geological period of the Paleozoic Era
groups Lobopodia and Radiodonta, Artiopoda (trilobites), Myriapoda and Pancrustacea. Found from Cambrian Stage 3, Radiodonta is characterised by a pair of
Cambrian
Group of arthropods
hypothesis — that Crustacea and Hexapoda form a monophyletic group, the Pancrustacea, to which the Myriapoda are the closest relatives — has support from
Uniramia
Species of mosquito
Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: Pancrustacea Class: Insecta Order: Diptera Family: Culicidae Genus: Culex Species:
London_Underground_mosquito
Class of crustaceans
Oligostraca and Multicrustacea, as the three main divisions of subphylum Pancrustacea, embracing the traditional crustaceans and the hexapods (including insects)
Remipedia
Many-legged arthropods with elongated bodies
Arthropoda Chelicerata inc. horseshoe crabs and spiders Mandibulata Myriapoda Chilopoda millipedes and allies Pancrustacea inc. crustaceans and insects
Centipede
Group of bees that feed on rotting meat
Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: Pancrustacea Class: Insecta Order: Hymenoptera Family: Apidae Genus: Trigona Subgenus:
Vulture_bee
Order of small freshwater animals
John Wiley and Sons. ISBN 978-0-471-35837-4. Evolution and Phylogeny of Pancrustacea: A Story of Scientific Method - 44. Notostraca and Allies (Calmanostraca)
Notostraca
Species of bee
Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: Pancrustacea Class: Insecta Order: Hymenoptera Family: Apidae Genus: Xylocopa Species:
Xylocopa_caerulea
Class of marine arthropods
Chelicerata Pycnogonida Euchelicerata Xiphosura and Arachnida Mandibulata Myriapoda Pancrustacea Crustacea and Hexapoda Best-supported position of Pycnogonida
Sea_spider
Subphylum of arthropods
Under the Mandibulata hypothesis, Myriapoda is the sister taxon to Pancrustacea, a group comprising the Crustacea and Hexapoda (insects and their close
Myriapoda
Extinct genus of arthropods
inconspicuous. It is hesitantly presumed to be an early offshoot under the clade Pancrustacea in original description. Other organisms, including Henningsmoenicaris
Goticaris
Extinct order of arthropods
the phylogenetic affinities of the enigmatic thylacocephalans within Pancrustacea". Proceedings of the Royal Society B: Biological Sciences. 292. Collette
Euthycarcinoidea
Species of butterfly
Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: Pancrustacea Class: Insecta Order: Lepidoptera Family: Papilionidae Genus: Papilio
Papilio_okinawensis
Extinct group of arthropods
Laville, T.; Bicknell, R. D. C. (2026). "Reassessing Concavicaris (Pancrustacea: Thylacocephala) from the Carboniferous of USA". Journal of Paleontology:
Thylacocephala
Species of flea
Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: Pancrustacea Class: Insecta Order: Siphonaptera Family: Hectopsyllidae Genus: Tunga
Tunga_penetrans
Species of true bug
Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: Pancrustacea Class: Insecta Order: Hemiptera Suborder: Heteroptera Family: Cimicidae
Cimex_hemipterus
Species of butterfly
Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: Pancrustacea Class: Insecta Order: Lepidoptera Family: Papilionidae Genus: Papilio
Papilio_castor
Species of insect
Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: Pancrustacea Class: Insecta Order: Hymenoptera Family: Apidae Genus: Apis Species:
Apis_laboriosa
Clade of arthropods
†Megacheira Pan-Chelicerata †Kiisortoqia †Bushizheia Antennulata †Artiopoda (including Trilobita) Mandibulata †Fuxianhuiida †Hymenocarina Myriapoda Pancrustacea
Deuteropoda
Clade of moths
Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: Pancrustacea Class: Insecta Order: Lepidoptera Infraorder: Heteroneura Clade: Eulepidoptera
Macroheterocera
Type of taxonomic group
(insects) are excluded. The modern clade that spans all of them is the Pancrustacea. One of the goals of modern taxonomy over the past fifty years has been
Paraphyly
Extinct order of arthropods
†Megacheira Pan-Chelicerata †Kiisortoqia †Bushizheia Antennulata †Artiopoda (including Trilobita) Mandibulata †Fuxianhuiida †Hymenocarina Myriapoda Pancrustacea
Fuxianhuiida
Species of termite
Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: Pancrustacea Class: Insecta Order: Blattodea Infraorder: Isoptera Family: Heterotermitidae
Heterotermes_aureus
Genus of wasps
Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: Pancrustacea Class: Insecta Order: Hymenoptera Family: Vespidae Subfamily: Polistinae
Synoeca
Extinct genus of arthropods
(2020-03-30). "Redescription of Americlus rankini (Woodward, 1868) (Pancrustacea: Cyclida: Americlidae) and interpretation of its systematic placement
Brittaniclus
Species of mantis shrimp
Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: Pancrustacea Class: Malacostraca Order: Stomatopoda Family: Gonodactylidae Genus:
Gonodactylus_smithii
Species of butterfly
Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: Pancrustacea Class: Insecta Order: Lepidoptera Family: Nymphalidae Genus: Euploea
Euploea_klugii
Common name for beetles that eat flour
Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: Pancrustacea Class: Insecta Order: Coleoptera Suborder: Polyphaga Infraorder: Cucujiformia
Flour_beetle
Species of small freshwater animal
Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: Pancrustacea Class: Branchiopoda Order: Notostraca Family: Triopsidae Genus: Triops
Triops_australiensis
Group of insects
Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: Pancrustacea Class: Insecta Order: Diptera Infraorder: Bibionomorpha Superfamily:
Fungus_gnat
Common name for two genera of wasps
Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: Pancrustacea Class: Insecta Order: Hymenoptera Family: Pompilidae Subfamily: Pepsinae
Tarantula_hawk
Species of beetle
Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: Pancrustacea Class: Insecta Order: Coleoptera Suborder: Polyphaga Infraorder: Elateriformia
Sternocera_aequisignata
Tribe of click beetles
Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: Pancrustacea Class: Insecta Order: Coleoptera Suborder: Polyphaga Infraorder: Elateriformia
Dendrometrini
Species of butterfly
Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: Pancrustacea Class: Insecta Order: Lepidoptera Family: Lycaenidae Genus: Pilodeudorix
Pilodeudorix_anetia
Tribe of beetles
Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: Pancrustacea Class: Insecta Order: Coleoptera Suborder: Polyphaga Infraorder: Cucujiformia
Dodecosini
Species of moth
Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: Pancrustacea Class: Insecta Order: Lepidoptera Family: Sphingidae Genus: Adhemarius
Adhemarius_jamaicensis
J. W. M.; Mychko, E. V. (2026). "A second enigmatic alsasuacaridid (Pancrustacea, Cyclida) from the Maastrichtian type area in The Netherlands (Late Cretaceous)"
2026 in arthropod paleontology
2026_in_arthropod_paleontology
Species of butterfly
Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: Pancrustacea Class: Insecta Order: Lepidoptera Family: Pieridae Genus: Gonepteryx
Gonepteryx_amintha
Genus of extinct arthropods
†Megacheira Pan-Chelicerata †Kiisortoqia †Bushizheia Antennulata †Artiopoda (including Trilobita) Mandibulata †Fuxianhuiida †Hymenocarina Myriapoda Pancrustacea
Isoxys
Species of moth
Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: Pancrustacea Class: Insecta Order: Lepidoptera Family: Tortricidae Genus: Cochylis
Cochylis_hollandana
Monotypic moth genus in family Drepanidae
Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: Pancrustacea Class: Insecta Order: Lepidoptera Family: Drepanidae Subfamily: Thyatirinae
Sugitaniella
Species of moth
Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: Pancrustacea Class: Insecta Order: Lepidoptera Family: Nepticulidae Genus: Acalyptris
Acalyptris_mariepsensis
Species of insect
Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: Pancrustacea Class: Insecta Order: Lepidoptera Family: Micropterigidae Genus: Micropterix
Micropterix_constantinella
Genus of flies
Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: Pancrustacea Class: Insecta Order: Diptera Family: Drosophilidae Genus: Samoaia Malloch
Samoaia
Genus of wasps
Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: Pancrustacea Class: Insecta Order: Hymenoptera Family: Bembicidae Subtribe: Gorytina
Liogorytes
Species of moth
Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: Pancrustacea Class: Insecta Order: Lepidoptera Superfamily: Noctuoidea Family: Erebidae
Apeplopoda_ochracea
Species of moth
Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: Pancrustacea Class: Insecta Order: Lepidoptera Family: Opostegidae Genus: Pseudopostega
Pseudopostega_mignonae
Species of butterfly
Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: Pancrustacea Class: Insecta Order: Lepidoptera Family: Lycaenidae Genus: Tajuria Species:
Tajuria_ister
Genus of moths
Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: Pancrustacea Class: Insecta Order: Lepidoptera Family: Pyralidae Subfamily: Epipaschiinae
Cacozelia
Genus of moths
Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: Pancrustacea Class: Insecta Order: Lepidoptera Family: Autostichidae Subfamily: Symmocinae
Orpecacantha
Species of moth
Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: Pancrustacea Class: Insecta Order: Lepidoptera Superfamily: Noctuoidea Family: Noctuidae
Dichagyris_singularis
Genus of moths
Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: Pancrustacea Class: Insecta Order: Lepidoptera Family: Crambidae Subfamily: Spilomelinae
Glycythyma
Species of praying mantis
Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: Pancrustacea Class: Insecta Order: Mantodea Family: Amorphoscelidae Genus: Amorphoscelis
Amorphoscelis_punctata
Species of butterfly
Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: Pancrustacea Class: Insecta Order: Lepidoptera Family: Nymphalidae Genus: Neptis Species:
Neptis_larseni
Superorder of crustaceans
A. Hegna, J. Luque, and J. A. Wolfe (2020). The fossil record of the Pancrustacea. In M. Thiel, G. C. B. Poore (eds.). Evolution and Biogeography of the
Eucarida
Superclass of crustaceans
Frederick R.; Koenemann, Stefan (31 March 2022). "Skaracarida". Evolution and Phylogeny of Pancrustacea: 92–97. doi:10.1093/oso/9780195365764.003.0007. v t e
Oligostraca
Species of wasp
Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: Pancrustacea Class: Insecta Order: Hymenoptera Family: Mymaridae Genus: Dicopomorpha
Dicopomorpha_echmepterygis
Species of butterfly
Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: Pancrustacea Class: Insecta Order: Lepidoptera Family: Lycaenidae Genus: Pentila Species:
Pentila_glagoessa
Species of butterfly
Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: Pancrustacea Class: Insecta Order: Lepidoptera Family: Lycaenidae Genus: Leptotes
Leptotes_adamsoni
Subfamily of wasps
Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: Pancrustacea Class: Insecta Order: Hymenoptera Family: Ichneumonidae Subfamily: Ophioninae
Ophioninae
Species of microscopic crustaceans
significance in the evolution of wings in insects, which evolved from within Pancrustacea. Abbreviations: at, antenna(e); cs, cephalic shield; le, lateral eye(s);
Yicaris
Beetles that emit a spray when threatened
Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: Pancrustacea Class: Insecta Order: Coleoptera Suborder: Adephaga Superfamily: Caraboidea
Bombardier_beetle
Species of insect
Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: Pancrustacea Class: Insecta Order: Diptera Family: Culicidae Genus: Anopheles Subgenus:
Anopheles_funestus
Species of insect endemic to Australia
Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: Pancrustacea Class: Insecta Order: Hymenoptera Family: Thynnidae Subfamily: Diamminae
Blue_ant
Species of beetle
Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: Pancrustacea Class: Insecta Order: Coleoptera Suborder: Polyphaga Infraorder: Scarabaeiformia
European_chafer
Genus of insects
Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: Pancrustacea Class: Insecta Order: Hemiptera Suborder: Heteroptera Family: Reduviidae
Rasahus
Species of butterfly
Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: Pancrustacea Class: Insecta Order: Lepidoptera Family: Lycaenidae Genus: Pseudophilotes
Pseudophilotes_sinaicus
Genus of praying mantises
Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: Pancrustacea Class: Insecta Order: Mantodea Family: Deroplatyidae Subfamily: Deroplatyinae
Deroplatys
Genus of wasps
Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: Pancrustacea Class: Insecta Order: Hymenoptera Family: Mymaridae Genus: Tinkerbella
Tinkerbella
Genus of planthoppers
Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: Pancrustacea Class: Insecta Order: Hemiptera Suborder: Auchenorrhyncha Infraorder:
Fulgora
Common name for several species of flies
Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: Pancrustacea Class: Insecta Order: Diptera Suborder: Nematocera Families See text
Midge
Species of ant
Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: Pancrustacea Class: Insecta Order: Hymenoptera Family: Formicidae Subfamily: Myrmicinae
Temnothorax_albipennis
Species of crustacean
Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: Pancrustacea Class: Malacostraca Order: Decapoda Suborder: Pleocyemata Infraorder:
Lysmata_debelius
Species of wasp
Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: Pancrustacea Class: Insecta Order: Hymenoptera Family: Mutillidae Genus: Dasymutilla
Dasymutilla_occidentalis
Genus of insects
Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: Pancrustacea Class: Insecta Order: Megaloptera Family: Corydalidae Subfamily: Corydalinae
Corydalus
Tribe of bees
Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: Pancrustacea Class: Insecta Order: Hymenoptera Family: Apidae Subfamily: Apinae Clade:
Bombini
Genus of beetles
Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: Pancrustacea Class: Insecta Order: Coleoptera Suborder: Polyphaga Infraorder: Cucujiformia
Coccoderus
Class of crustaceans
Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: Pancrustacea Superclass: Multicrustacea Class: Copepoda H. Milne-Edwards, 1840 Orders
Copepod
Genus of moths
Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: Pancrustacea Class: Insecta Order: Lepidoptera Family: Megalopygidae Genus: Macara
Macara_(moth)
Genus of beetles
Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: Pancrustacea Class: Insecta Order: Coleoptera Suborder: Polyphaga Infraorder: Cucujiformia
Lobopoda
Genus of bees
Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: Pancrustacea Class: Insecta Order: Hymenoptera Family: Apidae Tribe: Meliponini Genus:
Tetragona
Species of butterfly
Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: Pancrustacea Class: Insecta Order: Lepidoptera Family: Papilionidae Genus: Papilio
Papilio_polyctor
Species of stinging wasp
Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: Pancrustacea Class: Insecta Order: Hymenoptera Family: Vespidae Genus: Synoeca Species:
Synoeca_septentrionalis
Genus of beetles
Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: Pancrustacea Class: Insecta Order: Coleoptera Suborder: Polyphaga Infraorder: Cucujiformia
Thermistis
Species of moth
Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: Pancrustacea Class: Insecta Order: Lepidoptera Family: Geometridae Genus: Idaea Species:
Lophophleps_phoenicoptera
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