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Extinct group of arthropods
Thylacocephala (from the Greek θύλακος or thylakos, meaning "pouch", and κεφαλή or cephalon meaning "head") is an extinct group of mandibulate arthropods
Thylacocephala
Phylum of invertebrates with jointed exoskeletons
(Eurypterida) Apankura (Euthycarcinoidea) Trimerus (Artiopoda) Concavicaris (Thylacocephala) Cladogram after Liu et al., 2026: List of arthropod groups and genera
Arthropod
Superclass of crustaceans
†Genus Nothozoe Barrande, 1872 Incertae sedis †Order Cyclida †Order Thylacocephala Notes: Class Hexanauplia Oakley, Wolfe, Lindgren & Zaharof, 2013 was
Multicrustacea
Head section of an arthropod
The head of the Thylacocephala is also referred to as a cephalon: the cephalon is usually obscured by the carapace. Thylacocephala are a unique group
Cephalon_(arthropod_head)
Third and last period of the Mesozoic Era
sexual investment had the highest rates of extinction and turnover. Thylacocephala, a class of crustaceans, went extinct in the Late Cretaceous. The first
Cretaceous
Subphylum of arthropods
Oligostraca Ichthyostraca Mystacocarida Ostracoda Multicrustacea †Cyclida †Thylacocephala Copepoda Thecostraca Malacostraca Tantulocarida †Priscansermarinidae
Crustacean
Arthropods are invertebrate animals having an exoskeleton, a segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They
List_of_arthropod_orders
Clade of arthropods
†Martinssonia? †Papiliomaris? †Parioscorpio? †Rhyniognatha? †Skaracarida? †Tanazios? †Marrellomorpha? †Thylacocephala? †Oelandocarididae? †Musacaris?
Mandibulata
Possible extinct mandibulate arthropod
Devonian stem-arthropod Captopodus, and the somewhat enigmatic group Thylacocephala. Acheronauta has been assessed as a possible basal mandibulate, which
Acheronauta
Extinct order of crustaceans
the others being the freshwater-living Euthycarcinoida and the marine Thylacocephala. After Schweitzer, Mychko & Feldmann, 2020. Family Schraminidae Dzik
Cyclida
Extinct order of arthropods
Multicrustacea Copepoda Cirripedia (barnacles) †Cyclida (Americlidae) †Thylacocephala Malacostraca (shrimp, prawn, lobsters, mantis shrimp, crab, woodlice
Euthycarcinoidea
Extinct genus of bivalved invertebrate
recovered it as sister to Mandibulata in a clade comprising Acheronauta, Thylacocephala, and Captopodus. The other three hypotheses all independently produced
Papiliomaris
Extinct genus of arthropods
ravitis †Tanazios dokeron †Captopodus poschmanni †Acheronauta stimulapis †Thylacocephala †Occacaris oviformis Myriapoda †Ercaicunia multinodosa †Clypecaris pteroidea
Tanazios
Clade comprising all crustaceans and hexapods
Hexapoda) Multicrustacea Oligostraca †Hymenocarina †Phosphatocopina? †Thylacocephala? †Skaracarida? †Oelandocarididae? incertae sedis †Bredocaris †Dala †Ercaia
Pancrustacea
Lagerstätte in southwestern France
(5 October 2023). "Re-appraisal of thylacocephalans (Euarthropoda, Thylacocephala) from the Jurassic La Voulte-sur-Rhône Lagerstätte". European Journal
La Voulte-sur-Rhône (lagerstätte)
La_Voulte-sur-Rhône_(lagerstätte)
Extinct genus of arthropods
They noted these results are tentative. Below is the phylogeny of Thylacocephala as proposed by Schram (2014). ? indicates uncertain placement. Dames
Protozoea
Extinct genus of crustaceans
Scientific classification Kingdom: Animalia Phylum: Arthropoda Class: †Thylacocephala Order: †Conchyliocarida Genus: †Clausocaris Species: †C. lithographica
Clausocaris
Extinct genus of crustaceans
More recently, as in general for the group to which it belongs, the thylacocephala, the interpretation has shifted to a necrophagous or predatory organism
Ostenocaris
Silurian genus of arthropods
ravitis †Tanazios dokeron †Captopodus poschmanni †Acheronauta stimulapis †Thylacocephala †Occacaris oviformis Myriapoda †Ercaicunia multinodosa †Clypecaris pteroidea
Cascolus
Extinct genus of crustaceans
Scientific classification Kingdom: Animalia Phylum: Arthropoda Class: †Thylacocephala Order: †Conchyliocarida Family: †Dollocarididae Genus: †Dollocaris van
Dollocaris
Genus of Devonian arthropod
ravitis †Tanazios dokeron †Captopodus poschmanni †Acheronauta stimulapis †Thylacocephala †Occacaris oviformis Myriapoda †Ercaicunia multinodosa †Clypecaris pteroidea
Captopodus
Extinct genus of crustaceans
Scientific classification Kingdom: Animalia Phylum: Arthropoda Class: †Thylacocephala Order: †Concavicarida Genus: †Ainiktozoon Scourfield, 1937 Species:
Ainiktozoon
List of fauna present at the Mazon Creek fossil beds
Thomas; Haug, Joachim T.; Haug, Carolin (6 May 2021). "New species of Thylacocephala, Eodollocaris keithflinti n. gen., n. sp., from the Mazon Creek Lagerstätte
Paleofauna of the Mazon Creek fossil beds
Paleofauna_of_the_Mazon_Creek_fossil_beds
Extinct genus of crustaceans from Wisconsin
Scientific classification Kingdom: Animalia Phylum: Arthropoda Class: †Thylacocephala Family: †incertae sedis Genus: †Thylacares Species: †T. brandonensis
Thylacares
one'); Thylacoleo ('pouched lion'); Thylacosmilus ('pouched knife'); Thylacocephala ('pouch head') tri-: Pronunciation: /traɪ/. Origin: Ancient Greek τρία
List of commonly used taxonomic affixes
List_of_commonly_used_taxonomic_affixes
Bicknell, R. D. C. (2026). "Reassessing Concavicaris (Pancrustacea: Thylacocephala) from the Carboniferous of USA". Journal of Paleontology: 1–19. doi:10
2026 in arthropod paleontology
2026_in_arthropod_paleontology
Rock assemblage in Paris Canyon, Idaho, U.S.
A prawn Pemphix P. krumenackeri A glypheid pleocyemate crustacean Thylacocephala Gen. et sp. indet. A thylacocephalan Triassosculda T. ahyongi A mantis
Paris_biota
Geologic formation in Czech Republic
Rak, Štěpán; Budil, Petr; Fatka, Oldřich (2021). "Upper Ordovician Thylacocephala (Euarthropoda, Eucrustacea) from Bohemia indicate early ecological differentiation"
Letná_Formation
Mountain in Italy
Acanthinopus and (Palaeo)dusa have also been found. Of interest are the Thylacocephala, arthropods recently discovered taxonomically in the Besano fossiliferous
Cima_Rest
Geological formation in Italy and Switzerland
of the morphology and anatomy of Ostenocaris cypriformis (Crustacea, Thylacocephala)". Earth and Environmental Science Transactions of the Royal Society
Moltrasio_Formation
Arnaud Brayard; Sylvain Charbonnier (2021). "Review of Early Triassic Thylacocephala". Rivista Italiana di Paleontologia e Stratigrafia. 127 (1): 73–101
2019 in arthropod paleontology
2019_in_arthropod_paleontology
Paleontological site in Montana, USA
Schram, Frederick R. (2014-01-01). "Family level classification within Thylacocephala, with comments on their evolution and possible relationships". Crustaceana
Bear_Gulch_Limestone
thylacocephalans: A taphonomic case study of Austriocaris (Arthropoda, Thylacocephala) from the Fossil-Lagerstätte Polzberg (Reingraben shales, Carnian, Upper
List_of_lagerstätten
arthropod carapace, originally described as bivalved arthropod close to Thylacocephala, later reinterpreted as hurdiid radiodont close to Cambroraster, its
Paleobiota of the Maotianshan Shales
Paleobiota_of_the_Maotianshan_Shales
Ehiro; Osamu Sasaki; Harumasa Kano; Jun Nemoto; Hisayoshi Kato (2015). "Thylacocephala (Arthropoda) from the Lower Triassic of the South Kitakami Belt, Northeast
2015 in arthropod paleontology
2015_in_arthropod_paleontology
Lagerstätte Fossil site in Waukesha County, Wisconsin, U.S.
described as a separate genus from Venustulus. Thylacares Arthropoda Thylacocephala Six specimens Thylacares is a genus of thylacocephalan (a grouping of
Waukesha_Biota
Extinct genus of reptiles
Ji, C.; Tintori, A.; Jiang, D.; Motani, R. (2017). "New species of Thylacocephala (Arthropoda) from the Spathian (Lower Triassic) of Chaohu, Anhui Province
Cartorhynchus
Cretaceous (Albian to Cenomanian) El Doctor Formation Mexico A member of Thylacocephala (a group of arthropods of uncertain phylogenetic placement, possibly
2014 in arthropod paleontology
2014_in_arthropod_paleontology
PMC 11652639. PMID 39703206. Ji, C.; Tintori, A. (2024). "New Material of Thylacocephala from the Early Ladinian (Middle Triassic) of Northern Grigna (Lecco
2024 in arthropod paleontology
2024_in_arthropod_paleontology
Andrea Tintori; Dayong Jiang; Ryosuke Motani (2017). "New species of Thylacocephala (Arthropoda) from the Spathian (Lower Triassic) of Chaohu, Anhui Province
2017 in arthropod paleontology
2017_in_arthropod_paleontology
Jurassic Mayrocaris bucculata (Eucrustacea?, Thylacocephala) with comments on the tagmosis of Thylacocephala". Journal of Systematic Palaeontology. 19 (4):
2021 in arthropod paleontology
2021_in_arthropod_paleontology
A detailed study of Concavicarididae and Protozoeidae (?Crustacea, Thylacocephala) carapace micro-ornamentation". Journal of Systematic Palaeontology
2020 in arthropod paleontology
2020_in_arthropod_paleontology
Charbonnier, S. (2023). "Re-appraisal of thylacocephalans (Euarthropoda, Thylacocephala) from the Jurassic La Voulte-sur-Rhône Lagerstätte". European Journal
2023 in arthropod paleontology
2023_in_arthropod_paleontology
A detailed study of Concavicarididae and Protozoeidae (?Crustacea, Thylacocephala) carapace micro-ornamentation". Journal of Systematic Palaeontology
2018 in arthropod paleontology
2018_in_arthropod_paleontology
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