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PROTOPSEPHURUS

• Paddlefish

Family of fishes related to sturgeons

added in Hilton et al. (2023): genus † Protopsephurus Lu, 1994 (Early Cretaceous, China) species † Protopsephurus liui Lu, 1994 genus † Pugiopsephurus Hilton

Paddlefish



Paddlefish

• Protopsephurus

Extinct genus of fishes

Protopsephurus is an extinct genus of paddlefish containing the single species Protopsephurus liui, known from the Yixian, Jiufotang and Huajiying formations

Protopsephurus



Protopsephurus

• Acipenseriformes

Order of fishes

to 162 (195–137) million years ago. The oldest known paddlefish is Protopsephurus from the Early Cretaceous of China around 120 million years ago, while

Acipenseriformes



Acipenseriformes

• Chinese paddlefish

Extinct species of fish

over 190 million years ago. The oldest paddlefish fossil is that of Protopsephurus from the Early Cretaceous of China, dating to around 120 million years

Chinese paddlefish



Chinese_paddlefish

• American paddlefish

Species of freshwater fish

Lance; Jin, Fan; Yabumoto, Yoshitaka; Bemis, William E. (July 8, 2002). "Protopsephurus liui , a well-preserved primitive paddlefish (Acipenseriformes: Polyodontidae)

American paddlefish



American_paddlefish

• Syr Darya sturgeon

Species of fish

Polyodontidae †Crossopholis †Paleopsephurus †Parapsephurus Polyodon †Protopsephurus †Psephurus †Pugiopsephurus Acipenseridae Acipenser †Anchiacipenser †Boreiosturion

Syr Darya sturgeon



Syr_Darya_sturgeon

• Beluga (sturgeon)

Species of fish in the sturgeon family

Polyodontidae †Crossopholis †Paleopsephurus †Parapsephurus Polyodon †Protopsephurus †Psephurus †Pugiopsephurus Acipenseridae Acipenser †Anchiacipenser †Boreiosturion

Beluga (sturgeon)



Beluga_(sturgeon)

• Sturgeon

Ray-finned fish

(125-120 million years ago) or prior, as the earliest fossil paddlefish (Protopsephurus) is known from then. True sturgeons appear in the fossil record during

Sturgeon

Sturgeon

• Huso

Genus of fishes

Polyodontidae †Crossopholis †Paleopsephurus †Parapsephurus Polyodon †Protopsephurus †Psephurus †Pugiopsephurus Acipenseridae Acipenser †Anchiacipenser †Boreiosturion

Huso



Huso

• Adriatic sturgeon

Species of fish

Polyodontidae †Crossopholis †Paleopsephurus †Parapsephurus Polyodon †Protopsephurus †Psephurus †Pugiopsephurus Acipenseridae Acipenser †Anchiacipenser †Boreiosturion

Adriatic sturgeon



Adriatic_sturgeon

• Eochondrosteus

Extinct genus of fishes

Polyodontidae †Crossopholis †Paleopsephurus †Parapsephurus Polyodon †Protopsephurus †Psephurus †Pugiopsephurus Acipenseridae Acipenser †Anchiacipenser †Boreiosturion

Eochondrosteus

Eochondrosteus

• Siberian sturgeon

Species of fish

Polyodontidae †Crossopholis †Paleopsephurus †Parapsephurus Polyodon †Protopsephurus †Psephurus †Pugiopsephurus Acipenseridae Acipenser †Anchiacipenser †Boreiosturion

Siberian sturgeon



Siberian_sturgeon

• Shovelnose sturgeon

Species of fish

Polyodontidae †Crossopholis †Paleopsephurus †Parapsephurus Polyodon †Protopsephurus †Psephurus †Pugiopsephurus Acipenseridae Acipenser †Anchiacipenser †Boreiosturion

Shovelnose sturgeon



Shovelnose_sturgeon

• White sturgeon

Species of fish

Polyodontidae †Crossopholis †Paleopsephurus †Parapsephurus Polyodon †Protopsephurus †Psephurus †Pugiopsephurus Acipenseridae Acipenser †Anchiacipenser †Boreiosturion

White sturgeon



White_sturgeon

• Persian sturgeon

Species of fish

Polyodontidae †Crossopholis †Paleopsephurus †Parapsephurus Polyodon †Protopsephurus †Psephurus †Pugiopsephurus Acipenseridae Acipenser †Anchiacipenser †Boreiosturion

Persian sturgeon



Persian_sturgeon

• Caucasian sturgeon

Species of fish

Polyodontidae †Crossopholis †Paleopsephurus †Parapsephurus Polyodon †Protopsephurus †Psephurus †Pugiopsephurus Acipenseridae Acipenser †Anchiacipenser †Boreiosturion

Caucasian sturgeon

Caucasian_sturgeon

• Coccolepis

Extinct genus of fishes

Polyodontidae †Crossopholis †Paleopsephurus †Parapsephurus Polyodon †Protopsephurus †Psephurus †Pugiopsephurus Acipenseridae Acipenser †Anchiacipenser †Boreiosturion

Coccolepis



Coccolepis

• Parapsephurus

Extinct genus of fishes

which includes all polyodontids other than the most primitive genus Protopsephurus. Like Psephurus and other extinct paddlefish genera, the genus was likely

Parapsephurus



Parapsephurus

• Lycoptera

Extinct genus of fishes

within the Jehol Biota lake ecosystems, such as the primitive paddlefish Protopsephurus and the peipaiosteid (an extinct relative of sturgeons and paddlefish)

Lycoptera



Lycoptera

• Bastard sturgeon

Species of fish

Polyodontidae †Crossopholis †Paleopsephurus †Parapsephurus Polyodon †Protopsephurus †Psephurus †Pugiopsephurus Acipenseridae Acipenser †Anchiacipenser †Boreiosturion

Bastard sturgeon



Bastard_sturgeon

• Shortnose sturgeon

Species of fish

Polyodontidae †Crossopholis †Paleopsephurus †Parapsephurus Polyodon †Protopsephurus †Psephurus †Pugiopsephurus Acipenseridae Acipenser †Anchiacipenser †Boreiosturion

Shortnose sturgeon



Shortnose_sturgeon

• Crossopholis

Extinct genus of fishes

Lance; Jin, Fan; Yabumoto, Yoshitaka; Bemis, William E. (2002-07-08). "Protopsephurus liui , a well-preserved primitive paddlefish (Acipenseriformes: Polyodontidae)

Crossopholis



Crossopholis

• Priscosturion

Genus of fishes

Polyodontidae †Crossopholis †Paleopsephurus †Parapsephurus Polyodon †Protopsephurus †Psephurus †Pugiopsephurus Acipenseridae Acipenser †Anchiacipenser †Boreiosturion

Priscosturion



Priscosturion

• Paleopsephurus

Extinct genus of fishes

combination of characters which suggests that it is more derived than Protopsephurus, but is more basal than Crossopholis, Polyodon, or Psephurus. Relationships

Paleopsephurus

Paleopsephurus

• Kaluga (fish)

Species of sturgeon

Polyodontidae †Crossopholis †Paleopsephurus †Parapsephurus Polyodon †Protopsephurus †Psephurus †Pugiopsephurus Acipenseridae Acipenser †Anchiacipenser †Boreiosturion

Kaluga (fish)



Kaluga_(fish)

• Cylindracanthus

Extinct genus of fishes

Polyodontidae †Crossopholis †Paleopsephurus †Parapsephurus Polyodon †Protopsephurus †Psephurus †Pugiopsephurus Acipenseridae Acipenser †Anchiacipenser †Boreiosturion

Cylindracanthus



Cylindracanthus

• Chinese sturgeon

Species of fish

Polyodontidae †Crossopholis †Paleopsephurus †Parapsephurus Polyodon †Protopsephurus †Psephurus †Pugiopsephurus Acipenseridae Acipenser †Anchiacipenser †Boreiosturion

Chinese sturgeon

Chinese_sturgeon

• Amur sturgeon

Critically endangered species of fish

Polyodontidae †Crossopholis †Paleopsephurus †Parapsephurus Polyodon †Protopsephurus †Psephurus †Pugiopsephurus Acipenseridae Acipenser †Anchiacipenser †Boreiosturion

Amur sturgeon



Amur_sturgeon

• Acipenser

Genus of fishes

Polyodontidae †Crossopholis †Paleopsephurus †Parapsephurus Polyodon †Protopsephurus †Psephurus †Pugiopsephurus Acipenseridae Acipenser †Anchiacipenser †Boreiosturion

Acipenser



Acipenser

• Atlantic sturgeon

Species of fish

Polyodontidae †Crossopholis †Paleopsephurus †Parapsephurus Polyodon †Protopsephurus †Psephurus †Pugiopsephurus Acipenseridae Acipenser †Anchiacipenser †Boreiosturion

Atlantic sturgeon



Atlantic_sturgeon

• Russian sturgeon

Species of fish

Polyodontidae †Crossopholis †Paleopsephurus †Parapsephurus Polyodon †Protopsephurus †Psephurus †Pugiopsephurus Acipenseridae Acipenser †Anchiacipenser †Boreiosturion

Russian sturgeon



Russian_sturgeon

• Sinosturio

Genus of fishes

Polyodontidae †Crossopholis †Paleopsephurus †Parapsephurus Polyodon †Protopsephurus †Psephurus †Pugiopsephurus Acipenseridae Acipenser †Anchiacipenser †Boreiosturion

Sinosturio



Sinosturio

• Alabama sturgeon

Species of fish

Polyodontidae †Crossopholis †Paleopsephurus †Parapsephurus Polyodon †Protopsephurus †Psephurus †Pugiopsephurus Acipenseridae Acipenser †Anchiacipenser †Boreiosturion

Alabama sturgeon



Alabama_sturgeon

• Yixian Formation

Geological formation in China

Ordosemys (a prehistoric species of turtle) Palaeocambarus (a crawfish) Protopsephurus (an ancient species of paddlefish) Psittacosaurus preserved in combat

Yixian Formation



Yixian_Formation

• Amu Darya sturgeon

Species of fish

Polyodontidae †Crossopholis †Paleopsephurus †Parapsephurus Polyodon †Protopsephurus †Psephurus †Pugiopsephurus Acipenseridae Acipenser †Anchiacipenser †Boreiosturion

Amu Darya sturgeon



Amu_Darya_sturgeon

• Lake sturgeon

Species of fish

Polyodontidae †Crossopholis †Paleopsephurus †Parapsephurus Polyodon †Protopsephurus †Psephurus †Pugiopsephurus Acipenseridae Acipenser †Anchiacipenser †Boreiosturion

Lake sturgeon

Lake_sturgeon

• Gulf sturgeon

Species of fish

Polyodontidae †Crossopholis †Paleopsephurus †Parapsephurus Polyodon †Protopsephurus †Psephurus †Pugiopsephurus Acipenseridae Acipenser †Anchiacipenser †Boreiosturion

Gulf sturgeon



Gulf_sturgeon

• Dwarf sturgeon

Species of fish

Polyodontidae †Crossopholis †Paleopsephurus †Parapsephurus Polyodon †Protopsephurus †Psephurus †Pugiopsephurus Acipenseridae Acipenser †Anchiacipenser †Boreiosturion

Dwarf sturgeon



Dwarf_sturgeon

• Protoscaphirhynchus

Extinct genus of fishes

Polyodontidae †Crossopholis †Paleopsephurus †Parapsephurus Polyodon †Protopsephurus †Psephurus †Pugiopsephurus Acipenseridae Acipenser †Anchiacipenser †Boreiosturion

Protoscaphirhynchus

Protoscaphirhynchus

• Starry sturgeon

Species of fish

Polyodontidae †Crossopholis †Paleopsephurus †Parapsephurus Polyodon †Protopsephurus †Psephurus †Pugiopsephurus Acipenseridae Acipenser †Anchiacipenser †Boreiosturion

Starry sturgeon



Starry_sturgeon

• Pseudoscaphirhynchus

Genus of fishes

Polyodontidae †Crossopholis †Paleopsephurus †Parapsephurus Polyodon †Protopsephurus †Psephurus †Pugiopsephurus Acipenseridae Acipenser †Anchiacipenser †Boreiosturion

Pseudoscaphirhynchus



Pseudoscaphirhynchus

• European sea sturgeon

Species of fish

Polyodontidae †Crossopholis †Paleopsephurus †Parapsephurus Polyodon †Protopsephurus †Psephurus †Pugiopsephurus Acipenseridae Acipenser †Anchiacipenser †Boreiosturion

European sea sturgeon



European_sea_sturgeon

• Pallid sturgeon

Species of fish

Polyodontidae †Crossopholis †Paleopsephurus †Parapsephurus Polyodon †Protopsephurus †Psephurus †Pugiopsephurus Acipenseridae Acipenser †Anchiacipenser †Boreiosturion

Pallid sturgeon



Pallid_sturgeon

• Sakhalin sturgeon

Species of fish

Polyodontidae †Crossopholis †Paleopsephurus †Parapsephurus Polyodon †Protopsephurus †Psephurus †Pugiopsephurus Acipenseridae Acipenser †Anchiacipenser †Boreiosturion

Sakhalin sturgeon

Sakhalin_sturgeon

• Coccolepididae

Extinct family of ray-finned fishes

Polyodontidae †Crossopholis †Paleopsephurus †Parapsephurus Polyodon †Protopsephurus †Psephurus †Pugiopsephurus Acipenseridae Acipenser †Anchiacipenser †Boreiosturion

Coccolepididae



Coccolepididae

• Chondrosteidae

Extinct family of fishes

Polyodontidae †Crossopholis †Paleopsephurus †Parapsephurus Polyodon †Protopsephurus †Psephurus †Pugiopsephurus Acipenseridae Acipenser †Anchiacipenser †Boreiosturion

Chondrosteidae



Chondrosteidae

• Sterlet

Species of fish

Polyodontidae †Crossopholis †Paleopsephurus †Parapsephurus Polyodon †Protopsephurus †Psephurus †Pugiopsephurus Acipenseridae Acipenser †Anchiacipenser †Boreiosturion

Sterlet



Sterlet

• Scaphirhynchus

Genus of fishes

Polyodontidae †Crossopholis †Paleopsephurus †Parapsephurus Polyodon †Protopsephurus †Psephurus †Pugiopsephurus Acipenseridae Acipenser †Anchiacipenser †Boreiosturion

Scaphirhynchus



Scaphirhynchus

• Strongylosteus

Extinct genus of fishes

Polyodontidae †Crossopholis †Paleopsephurus †Parapsephurus Polyodon †Protopsephurus †Psephurus †Pugiopsephurus Acipenseridae Acipenser †Anchiacipenser †Boreiosturion

Strongylosteus



Strongylosteus

• Green sturgeon

Species of fish

Polyodontidae †Crossopholis †Paleopsephurus †Parapsephurus Polyodon †Protopsephurus †Psephurus †Pugiopsephurus Acipenseridae Acipenser †Anchiacipenser †Boreiosturion

Green sturgeon



Green_sturgeon

• Feathered Dinosaurs of China

Book by Gregory Wenzel

Predatory Fish (seen in the distance, hidden under some Water Lilies) Protopsephurus Lycoptera Hyphalosaurus Unidentified Cicadas (seen on page 20) Beipiaosaurus

Feathered Dinosaurs of China

Feathered_Dinosaurs_of_China

• Gyrosteus

Extinct genus of fishes

Polyodontidae †Crossopholis †Paleopsephurus †Parapsephurus Polyodon †Protopsephurus †Psephurus †Pugiopsephurus Acipenseridae Acipenser †Anchiacipenser †Boreiosturion

Gyrosteus

Gyrosteus

• Chondrostei

Subclass of fishes

Polyodontidae †Crossopholis †Paleopsephurus †Parapsephurus Polyodon †Protopsephurus †Psephurus †Pugiopsephurus Acipenseridae Acipenser †Anchiacipenser †Boreiosturion

Chondrostei

Chondrostei

• Chondrosteus

Extinct genus of fishes

Polyodontidae †Crossopholis †Paleopsephurus †Parapsephurus Polyodon †Protopsephurus †Psephurus †Pugiopsephurus Acipenseridae Acipenser †Anchiacipenser †Boreiosturion

Chondrosteus



Chondrosteus

• Jiufotang Formation

Geological formation in Liaoning, China

Lycoptera L. davidi An osteoglossomorph Peipiaosteus P. pani A fish related to sturgeons Protopsephurus P. liui A paddlefish Sinamia S. zdanskyi An amiiform

Jiufotang Formation

Jiufotang_Formation

• Birgeria

Extinct genus of fishes

Polyodontidae †Crossopholis †Paleopsephurus †Parapsephurus Polyodon †Protopsephurus †Psephurus †Pugiopsephurus Acipenseridae Acipenser †Anchiacipenser †Boreiosturion

Birgeria



Birgeria

• Paleobiota of the Yixian Formation

Geological formation in China

Protopsephurus

Paleobiota of the Yixian Formation



Paleobiota_of_the_Yixian_Formation

• Guildayichthyidae

Extinct order of ray-finned fishes

Polyodontidae †Crossopholis †Paleopsephurus †Parapsephurus Polyodon †Protopsephurus †Psephurus †Pugiopsephurus Acipenseridae Acipenser †Anchiacipenser †Boreiosturion

Guildayichthyidae



Guildayichthyidae

• Peipiaosteus

Extinct genus of fishes

Polyodontidae †Crossopholis †Paleopsephurus †Parapsephurus Polyodon †Protopsephurus †Psephurus †Pugiopsephurus Acipenseridae Acipenser †Anchiacipenser †Boreiosturion

Peipiaosteus



Peipiaosteus

• Huajiying Formation

Geologic formation in China

tokunagai Peipiaosteus P. fengningensis A fish related to sturgeons Protopsephurus P. liui Also present in the Jiufotang & Yixian Formations A paddlefish

Huajiying Formation

Huajiying_Formation

• Morrolepis

Extinct genus of ray-finned fishes

Polyodontidae †Crossopholis †Paleopsephurus †Parapsephurus Polyodon †Protopsephurus †Psephurus †Pugiopsephurus Acipenseridae Acipenser †Anchiacipenser †Boreiosturion

Morrolepis



Morrolepis

• Jianianhualong

Extinct genus of dinosaurs

including Lycoptera (of which L. davidi is numerous), Peipiaosteus, and Protopsephurus. Environmentally, Jianianhualong lived in a lacustrine area, as evinced

Jianianhualong



Jianianhualong

• Geology of China

Kong Science Museum, Ductina vietnamica from the Devonian (Hunan), Protopsephurus fossil specimen excavated in Liaoning and the holotype of Microraptor

Geology of China



Geology_of_China

• Stichopterus

Extinct genus of fishes

Polyodontidae †Crossopholis †Paleopsephurus †Parapsephurus Polyodon †Protopsephurus †Psephurus †Pugiopsephurus Acipenseridae Acipenser †Anchiacipenser †Boreiosturion

Stichopterus

Stichopterus

• Plesiococcolepis

Extinct genus of ray-finned fishes

Polyodontidae †Crossopholis †Paleopsephurus †Parapsephurus Polyodon †Protopsephurus †Psephurus †Pugiopsephurus Acipenseridae Acipenser †Anchiacipenser †Boreiosturion

Plesiococcolepis

Plesiococcolepis

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PROTOPSEPHURUS

Online names & meanings

• Srinivasulu

Boy/Male

Hindu

Srinivasulu

Lord venkateswara

• Bourey

Boy/Male

Cambodian

Bourey

County.

• Gibbar

Boy/Male

Biblical

Gibbar

Strong; manly.

• Guni

Boy/Male

Biblical

Guni

A garden, a covering.

• LÓRÁNT

Male

Hungarian

LÓRÁNT

Hungarian form of Norman French Roland, LÓRÁNT means "famous land."

• Kuntibhoja

Boy/Male

Hindu, Indian, Traditional

Kuntibhoja

Bearer of Spears

• Heeter

Surname or Lastname

North German

Heeter

North German : habitational name for someone from Heeten in the Netherlands near Deventer.English : unexplained; perhaps a variant of Hayter. Compare Heater.

• Murtada |

Boy/Male

Muslim

Murtada |

Satisfied, Contented, Pleased, Chosen

• Prithiv

Boy/Male

Hindu

Prithiv

The Sun

• SHALOTT

Female

Arthurian

SHALOTT

, shallot (the onion); or, Alclut, the name of the rock of Dumbarton.

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PROTOPSEPHURUS

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PROTOPSEPHURUS

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PROTOPSEPHURUS