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First cloned mammal (1996–2003)
Dolly (5 July 1996 – 14 February 2003) was a female Finn-Dorset sheep and the first mammal that was cloned from an adult somatic cell. She was cloned
Dolly_(sheep)
American singer-songwriter (1946–2026)
Dolly Rebecca Parton Dean (née Parton; January 19, 1946 – August 25, 2026) was an American singer-songwriter, actress, philanthropist, and businesswoman
Dolly_Parton
has not continued since then. Dolly (sheep) Pharming (genetics) Genetically modified organisms "Tracy, a transgenic sheep, Scotland, 1999". www.sciencemuseum
Tracy_(sheep)
Topics referred to by the same term
Look up dolly in Wiktionary, the free dictionary. Dolly is a synonym for a doll Dolly may also refer to: Dolly (tool), a portable anvil Dolly pot, also
Dolly
Animal husbandry
Sheep farming or sheep husbandry is the raising and breeding of domestic sheep. It is a branch of animal husbandry. Sheep are raised principally for their
Sheep_farming
British biologist and university professor (1954–2012)
world by successfully cloning a sheep from adult mammary cells. Dolly, a Finn Dorset sheep named after the singer Dolly Parton, was born in 1996 and lived
Keith_Campbell_(biologist)
First cloned animal in Turkey
2011. Oyah had died from a progressive lung disease on 16 April 2012. Dolly (sheep) List of cloned animals "Oyalı sizlere ömür". HaberTürk. 18 April 2012
Oyalı_(sheep)
2022 book by Chuck Klosterman
J. Simpson 1996 Russian presidential election Waco siege Cloning of Dolly (sheep) 1994–95 Major League Baseball strike Rodney King riots Oklahoma City
The_Nineties_(book)
Breed of sheep
of the Finnsheep's prolificacy and the Dorset's meat conformation. Dolly the sheep, first mammal to be cloned from an adult somatic cell, was a Finn Dorset
Finn_Dorset
Species of domesticated mammal
Sheep (pl.: sheep) or domestic sheep (Ovis aries) are a domesticated ruminant mammal typically kept as livestock. Although the term sheep can apply to
Sheep
domestic sheep, were the first mammals to have been successfully cloned from differentiated cells. They are not to be confused with Dolly the sheep which
Megan_and_Morag
U.S. wilderness area in West Virginia
The Dolly Sods Wilderness (DSW, originally simply Dolly Sods) is a U.S. Wilderness Area in the Allegheny Mountains of eastern West Virginia and is part
Dolly_Sods_Wilderness
Pair of cloned sheep
transgenic animals at the same time. This is not to be confused with Dolly the Sheep, the first animal to be successfully cloned from an adult somatic cell
Polly_and_Molly
British embryologist (1944–2023)
Archived from the original on 15 June 2009. Retrieved 16 April 2009. "Dolly the sheep creator knighted". BBC. 29 December 2007. Retrieved 30 December 2007
Ian_Wilmut
Cellular process of loss of specialization
as well as the possible risks can be found here [1] . Dolly (sheep), a female Finn-Dorset sheep and the first mammal that was cloned from an adult somatic
Dedifferentiation
Process of producing individual organisms with identical genomes
viable embryo is created from a somatic cell and an egg cell. In 1996, Dolly the sheep achieved notoriety for being the first mammal cloned from an adult
Cloning
Philippine film studio, subsidiary of Star Cinema
April 27, 2023. "Kathryn Bernardo to star in 3 films, including one with Dolly de Leon". Rappler. March 28, 2023. Black Sheep Productions on Facebook
Black_Sheep_Productions
Scottish animal scientist (1948–2015)
known for her contribution as part of the pioneering team who cloned Dolly The Sheep. Ritchie was born in Edinburgh, Scotland on 29 March 1948 to Helen
Marjorie_Ritchie
Chief Veterinary Officer of the United Kingdom from 2004 to 2007
original on 6 September 2008. Retrieved 10 November 2007. "Honour for Dolly sheep scientist". BBC News. 29 December 2007. Retrieved 5 January 2008. "Profile
Debby_Reynolds
2014 decision of the United States Court of Appeals for the Federal Circuit
rejecting a patent for a cloned sheep known as "Dolly the Sheep"— the first mammal ever cloned from an adult somatic cell. Dolly was cloned in 1996 by Ian Wilmut
In re Roslin Institute (Edinburgh)
In_re_Roslin_Institute_(Edinburgh)
Scottish animal sciences research institute
Biological Sciences Research Council. It is best known for creating Dolly the sheep in 1996, the first mammal to be successfully cloned from an adult cell
Roslin_Institute
World's first cloned primates (born 2017)
nuclear transfer (SCNT), the same cloning technique that produced Dolly the sheep in 1996. They are the first cloned primates produced by this technique
Zhong_Zhong_and_Hua_Hua
British legal scholar
A finding was that UK school students had been misinformed as to how Dolly (sheep) had been cloned. In January 2023, Pattinson published Law at the Frontiers
Shaun_Pattinson
Fictional character from Emmerdale
second marriage, Matt leaves the village to run a sheep farm in Norfolk, but later remarries Dolly after she and Sam join him. Matt was mentioned after
Matt_Skilbeck
1973 American film
introducing asexual reproduction via cloning to pop culture; it predates Dolly the Sheep by 23 years. At the Pacific Institute of Technology, nuclear scientist
The_Clones_(film)
Form of cloning
a cloned organism develops to term in rare instances. This is how Dolly the Sheep and many other species were cloned. Cows are commonly cloned to select
Nuclear_transfer
2002 video-opera by Steve Reich
the Hindenburg, nuclear testings on Bikini Atoll, and the cloning of Dolly the sheep (drawing connections between genetic engineering and artificial intelligence)
Three_Tales_(opera)
Commercial cloning of animals
extinct animals. The practice was first demonstrated in 1996 with Dolly the sheep. Moving or copying all (or nearly all) genes from one animal to form
Commercial_animal_cloning
English actor, director and writer
his performances in the West End revivals of Fiddler on the Roof, Hello, Dolly! and The Producers. Nyman's first noteworthy performance was in 2000 as
Andy_Nyman
Type of pluripotent blastocystic stem cell
variety of cloning techniques. The Dolly experiment was performed by obtaining the mammalian udder cells from a sheep (Dolly) and differentiating these cells
Embryonic_stem_cell
Prince and Princess of Wales (Charles and Diana) and the birth of Dolly the sheep. Monarch – Elizabeth II Prime Minister – John Major (Conservative)
1996_in_the_United_Kingdom
1923 film
Clifford S. Efelt, directed by Otis B. Thayer, and starring Edmund Cobb, Dolly Dale, Helen Hayes and Frank Gallagher. The film was shot in Colorado by
Riders of the Range (1923 film)
Riders_of_the_Range_(1923_film)
2000 studio album by Buck-Tick
comes from lyrics to "Cyborg Dolly: Sora-mimi: Phantom", which also mentions cloning sheep and is named after the famous Dolly. It reached number eleven
One_Life,_One_Death
1970 compilation album by various artists
"Entropy" – Quatermass "Black Sheep Of The Family" – Quatermass "Grass" – The Pretty Things "Salisbury Plain" – Shirley & Dolly Collins "Embryo" – Pink Floyd
Picnic – A Breath of Fresh Air
Picnic_–_A_Breath_of_Fresh_Air
Dutch comic series
unnamed in the comics, but received names in the animated series. They are Dolly the tortoise, Shampoo the gorilla and Saffie the dog. In the English dub
Boes
Internet meme
(called Holly Dolly) dancing to the animation which is displayed (flipped horizontally) in the background. The animation is marketed as the "Dolly Song", and
Loituma_Girl
large mammal was a sheep by Steen Willadsen in 1984. However, the cloning was done from early embryonic cells, while the sheep Dolly in 1996 was cloned
List_of_cloned_animals
Danish scientist
embryos to clone sheep by nuclear transfer. The procedure he developed was essentially identical to the one used to create Dolly the sheep, although nuclei
Steen_Willadsen
Method of creating a cloned embryo by replacing the egg nucleus with a body cell nucleus
It is used in both therapeutic and reproductive cloning. In 1996, Dolly the sheep became famous for being the first successful case of the reproductive
Somatic_cell_nuclear_transfer
said there would be no danger to the eater if Rubis was consumed. Dolly the sheep Her name is sometimes translated as Ruby in English reports Gonzalez
Rubis_(sheep)
San Francisco-based software company
Dolly after Dolly the sheep, that developers could use to create custom chatbots. Dolly has only 6 billion parameters. Databricks claimed that Dolly had
Databricks
National Park located in Yukon, Canada
lemmings, Dall sheep, gyrfalcons, muskoxen, and wolverines. The area around Sheep Creek is the northernmost extension of Dall sheep habitat in Canada
Ivvavik_National_Park
English writer
biological science. The original findings detailing the cloning of Dolly the sheep by Ian Wilmut and his team at the Roslin Institute in Scotland were
Robin_McKie
English geneticist
development of the new industry of bio-farming. In 1996 the cloning of Dolly the sheep at Roslin received wide attention both in the public domain and in
Grahame_Bulfield
Cannon, Lady Dorothy "Dolly" Beatrice Louisa Hale is one of Georgina's closest friends and a fellow "Bright Young Thing". Lady Dolly is the daughter of the
List of Upstairs, Downstairs (1971 TV series) characters
List_of_Upstairs,_Downstairs_(1971_TV_series)_characters
Overview of and topical guide to biochemistry
create human insulin. Cloning – Dolly the sheep was the first mammal ever cloned from adult animal cells. The cloned sheep was, of course, genetically identical
Outline_of_biochemistry
Animal cloning company
competitor is somatic cell nuclear transfer, as which was used for cloning Dolly the Sheep. Viagen was founded in Austin, Texas in 2002. Viagen began by offering
Viagen
2025). "Mokster Films Shops Taiwanese Roto-Animated Movie 'Welcome to Dolly's House' at Cannes". Animation Magazine. Retrieved July 15, 2026. Wong, Silvia
List of animated feature films of 2026
List_of_animated_feature_films_of_2026
Kiersey Clemons, Ken Marino, Claudia Sulewski, Simon Helberg, Liv Hewson Dolly Independent Film Company / Shudder Rod Blackhurst (director/screenplay);
List of American films of 2026
List_of_American_films_of_2026
Trailer vehicle without a front axle
supported by a tractor unit, or a detachable front-axle assembly known as a dolly, or the tail of another trailer. The semi-trailer's weight is semi-supported
Semi-trailer
American actress (1924–2019)
by bombardiers of the US Army Air Forces. In 1947 she played the part of Dolly in the Alexander Hall-directed musical fantasy Down to Earth, which again
Kathleen_O'Malley
The same technique, using somatic cells, was later used to create Dolly the Sheep. Their experiment was the first successful nuclear transplantation
Robert_Briggs_(scientist)
orbits Jupiter, studying the planet and its moons extensively. 1996 Dolly the sheep is cloned. Construction starts on the International Space Station.
1990s in science and technology
1990s_in_science_and_technology
American comedian, actor (born 1988)
film 2019 Late Night Chris Reynolds Nisha Ganatra 2019 Good Posture Sol Dolly Wells 2020 Save Yourselves! Blake Alex Huston Fischer Eleanor Wilson 2020
John_Early_(comedian)
histologically similar to human adenocarcinoma in situ of the lung. Dolly the sheep, the first mammal successfully cloned from an adult somatic cell, was
Ovine pulmonary adenocarcinoma
Ovine_pulmonary_adenocarcinoma
Filipino and American actress (born 1998)
pigeonholed as motivation to build a versatile body of work. She cites actress Dolly de Leon as an inspiration for her acting pursuits after the latter received
Liza_Soberano
American game show
"Wednesday TV Ratings 8/26/26: Jeopardy! Masters Jumps on Second Night, Dolly Parton Specials Strong on NBC and CW, Big Brother Leads". The TV Ratings
Nation's_Dumbest
19th-century American criminal
including Billy Morton, Frank Baker, Jim McDaniels, Buffalo Bill Spawn, Dolly Graham, Tom Hill, Bob Martin, Nicholas Provencio, and Manuel Segovia. Although
Jesse_Evans
2022 season of an American television series
William R. Moses as Yan Scott Speiser as Verell Elyse Levesque as Wenda Dolly Parton as a simulation of herself Giorgia Whigham as Lysella Halston Sage
The_Orville_season_3
Type of knot
variations of sheep shank using either a simple half hitch or a double turn self crossing half hitch or a triple turn self crossing half hitch. A sheep shank
Trucker's_hitch
Village in Midlothian, Scotland
Roslin Institute, a biological research establishment, where in 1996 Dolly the sheep became the first animal to be cloned from an adult somatic cell. She
Roslin,_Midlothian
American actress (born 1958)
in 1979. She made her New York City Opera debut in 1982's Candide (Pink Sheep) and remained on the guest artist roster through 1989. Other NYCO credits
Ivy_Austin
American actress (1919–2000)
as Thora Flagg in "The Mutton Puncher" (both 1957) The Californians as Dolly Dawson in "The Regulators" (1957) Maverick in the episodes "The Quick and
Marie_Windsor
Hannah Jane Parkinson (2017-10-23), "From Félicette the space cat to Dolly the sheep – which animals should be given a statue?", The Guardian, retrieved
List of individual hippopotamids
List_of_individual_hippopotamids
production A Very Good Girl September 27 Petersen Vargas Kathryn Bernardo and Dolly de Leon Mystery Film, Dark comedy Rewind December 25 Mae Cruz-Alviar Dingdong
List of films produced and released by Star Cinema
List_of_films_produced_and_released_by_Star_Cinema
Irish animated children's television series
features a subplot featuring a sheep named Wiley (voiced by Mel Brooks), who lives at Raloo Farm with his flock. As the only sheep in the flock who can talk
Jakers! The Adventures of Piggley Winks
Jakers!_The_Adventures_of_Piggley_Winks
Topics referred to by the same term
Campbell (biologist) (1954–2012), British biologist, led team that cloned Dolly the sheep Keith Campbell (cricketer) (born 1943), New Zealand cricketer Keith
Keith_Campbell
American actor (1902–1971)
28, 2016. The Broadway League. "Hello, Dolly! – IBDB: The official source for Broadway Information". Hello, Dolly! Playbill. Retrieved October 12, 2017
David_Burns_(actor)
dies at age 52. August 25: Dolly Parton, American singer-songwriter, actress, philanthropist, and businesswoman (voice of Dolly Gnome in Gnomeo and Juliet
2026_in_animation
Topics referred to by the same term
Edinburgh, Scotland Rosslyn Chapel Roslin Castle Roslin Institute, where Dolly the Sheep was cloned Battle of Roslin, 1303 Barony of Roslin Roslin House, the
Roslin
British animal geneticist
scientists responsible for laying the groundwork for the creation of Dolly the Sheep. The Roger Land Building within the University of Edinburgh's King's
Roger_Burton_Land
use as evidence Kadhal Reset Repeat During the credits, Aditi voicing. Dolly The metal screen door opens and Officer Lee along with another officer and
List of films with post-credits scenes (2020s)
List_of_films_with_post-credits_scenes_(2020s)
American actress and comedienne (b. 1937)
as a standby for the lead in the original Broadway production of Hello, Dolly! One year later, Worley created her own nightclub act in Greenwich Village
Jo_Anne_Worley
1876–80 gang in New Mexico, United States
breaking away, he brought along with him Billy Morton, Frank Baker, Tom Hill, Dolly Graham, George Davis, Jim McDaniels, Buffalo Bill Spawn, Bob Martin, Manuel
Jesse_Evans_Gang
Japanese fashion subculture
apparel pieces vary and manba, as they would replace the colors worn with dolly pastels, pink lace and sundresses that are frilled. Romanba gyarus instead
Gyaru
Bacterial infection characterized by muscle spasms
840C. doi:10.1016/0006-291X(85)90123-8. PMID 3994725. Foran P, Shone CC, Dolly JO (December 1994). "Differences in the protease activities of tetanus and
Tetanus
2026 British TV series
2026-05-04 – via www.bbc.co.uk. Tiny Tunes - Series 1: 18. Miss Polly Had a Dolly. Retrieved 2026-05-04 – via www.bbc.co.uk. Tiny Tunes - Series 1: 19. Oranges
Tiny_Tunes_(TV_series)
Annual celebration of the harvest season
heard on each day of the harvest. A corn dolly was made from the last sheaf of corn harvested. The corn dolly often had a place of honour at the banquet
Harvest_festival
British actress and comedian (born 1966)
alongside Sally Phillips and Nick Hamson, of the production company Captain Dolly. In 2026, she played the role of Bea Pollard in the BBC soap opera, EastEnders
Ronni_Ancona
2017 film by Denis Villeneuve
with Zeiss Master Prime lenses, assisted with an attached crane arm or a dolly. The filmmakers conducted tests with an Alexa 65 camera but preferred the
Blade_Runner_2049
Dogs Go to Heaven Gabriel in Constantine The Escort in Heaven Can Wait Dolly Parton in Unlikely Angel The fallen angels in Noah Satan in Wreck-It Ralph
List_of_stock_characters
Taiwanese-American neuroscientist (born 1948)
using somatic cell nuclear transfer (the technique used to create Dolly the sheep) rather than embryo twinning. According to Poo, the principal significance
Mu-ming_Poo
British television period-drama
episodes; all series). Jameson is Bolam's wife off screen. Madelaine Newton as Dolly Mather/Ford/Seaton (30 episodes; series 1–3) Basil Henson as Sir Horatio
When_the_Boat_Comes_In
British evolutionary biologist
W D Hamilton. Vol. 25, no. 3. London Review of Books. "Slate". Dolly the Cloned Sheep. Slate. 2 March 1997. Alfred Russel Wallace; Andrew Berry; Stephen
Andrew_Berry_(biologist)
English molecular biologist
Ian Wilmut's team at Roslin to clone a sheep, Dolly (1996), the result of transplanted DNA of an adult sheep to an unfertilized egg cell. He was appointed
Anthony_John_Clark
American television director and actor (born 1938)
run of Hello, Dolly!, he stepped into the role of Cornelius Hackl created by Charles Nelson Reilly, and he also appeared in the plays Sheep on the Runway
Will_Mackenzie
American veteran of the War of 1812
the War of 1812, namer of Cumberland, Maine, first person to bring merino sheep to Maine, a ship captain, and the first treasurer of Cumberland. Sturdivant
Ephraim_Sturdivant
STRAIGHT-LINE WINDS, AND FLOODING DR-1779-NE 2008 Severe Storm TX HURRICANE DOLLY DR-1780-TX 2008 Hurricane ID FLOODING DR-1781-ID 2008 Flood NH SEVERE STORMS
List of FEMA disaster and other emergency declarations
List_of_FEMA_disaster_and_other_emergency_declarations
Museum in Edinburgh, Scotland
on display. One of the more notable exhibits is the stuffed body of Dolly the sheep, the first successful cloning of a mammal from an adult cell. Other
National_Museum_of_Scotland
2000 film directed by Roger Spottiswoode
a total budget of $82 million. Following the successful cloning of Dolly the sheep in the 1990s, artificial cloning of animals has become commonplace
The_6th_Day
2008 British TV series or programme
Damer as Lenny; Sam Phillips as Jamie, the bellboy; and Tanya Franks as Dolly and Sheila Bernette as Mrs. Poshington, a guest who never leaves (and who
Hotel_Trubble
Polly Had a Dolly" is also referred to as; 'Miss Molly Had a Dolly', 'Miss Polly', 'Miss Polly had a little dolly', 'Miss Polly had a dolly who was sick
List_of_nursery_rhymes
American actor (born 1992)
singers Evan and Jaron Lowenstein. Lerman has stated that he is a "black sheep" in his household because he is an actor, while most of his relatives work
Logan_Lerman
Film by Ron Underwood
eruption shots. After the creature emerged, it could be attached to a rolling dolly rig suspended from a crane for shots in which it moved across the set, including
Tremors_(1990_film)
American rock band
Animal.[citation needed] On November 30, 2018, Dorothy appeared along with Dolly Parton on the soundtrack album for Parton's film "Dumplin'". On November
Dorothy_(band)
American-British rock musician (born 1950)
2013. Coon, Caroline (July 20, 1974). "Suzi Quatro & Olivia Newton-John: Dolly Mixture". Melody Maker. (subscription required) "Previous Honorees". Sherocksawards
Suzi_Quatro
American science fiction comedy drama television series (2017–2022)
ethical dimensions of gender reassignment surgery and its cameo featuring Dolly Parton. He also gave a favorable appraisal of the time travel episode "Twice
The_Orville
2011 film by George Nolfi
fit the conditions of the day. The visual plan for the film was to use a dolly or crane to keep camera movements smooth and employ a more formal style
The_Adjustment_Bureau
Fictional character from My Little Pony
way." Voice actress Ashleigh Ball has cited country singers Miley Cyrus, Dolly Parton, and Reba McEntire as the main influences on Applejack's voice, which
Applejack_(My_Little_Pony)
English comedian and writer (born 1967)
toothbrush moustache and moving onto the hoodie, Flag of England and Dolly the Sheep. An Edinburgh special about the See-you-Jimmy hat was broadcast in
Richard_Herring
travel, tourism, insurance
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travel, tourism, insurance