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Extinct species of canine mammal
The dire wolf (Aenocyon dirus /aɪˈnɒkaɪ.ɒn ˈdaɪrəs/ ) is an extinct species of canine which was native to the Americas during the Late Pleistocene and
Dire_wolf
Bioengineering project with the goal of replicating an extinct species
Colossal Biosciences dire wolf project is a project by Colossal Biosciences with the goal of replicating the phenotype of the extinct dire wolf (Aenocyon dirus)
Colossal Biosciences dire wolf project
Colossal_Biosciences_dire_wolf_project
Topics referred to by the same term
A dire wolf is an extinct canine. Dire wolf or direwolf may also refer to: Direwolf (Game of Thrones), a fictional creature in George R. R. Martin's A
Dire_wolf_(disambiguation)
1970 song by Grateful Dead
"Dire Wolf" is a ballad by the Grateful Dead, released as the third track on their 1970 album Workingman's Dead. The lyrics were written by Robert Hunter
Dire_Wolf_(song)
American biotechnology company
extinct animals, including the woolly mammoth, the Tasmanian tiger, the dire wolf, the dodo, the moa and the bluebuck. In 2023, it stated that it wants
Colossal_Biosciences
Process of re-creating an extinct organism
been attempts to recreate the dire wolf (Aenocyon dirus) in modern times. The first is a project called the Dire Wolf Project, a project begun in 1988
De-extinction
lineages evolved in isolation from the dire wolf lineage. The study proposes an early origin of the dire wolf lineage in the Americas, and that this geographic
Evolution_of_the_wolf
Extinct North American wolf
to its skull size. In comparison with the Beringian wolf, the more southerly occurring dire wolf (Aenocyon dirus) was the same size but heavier and with
Beringian_wolf
2020 board game by Paul Dennen
Imperium is a 2020 board game designed by Paul Dennen and published by Dire Wolf Digital. In the board game, which is set in Frank Herbert's Dune universe
Dune:_Imperium
Extinct species of carnivore
Early to Middle Pleistocene epoch. It is proposed as the ancestor of the dire wolf, one of the most famous prehistoric carnivores in North America, and likely
Armbruster's_wolf
Species of canine
showed an increase in tooth breakage similar to that seen in the extinct dire wolf. This suggests they either often processed carcasses, or that they competed
Wolf
Extinct species of canid
the dire wolf. The sequences indicate the dire wolf to be a highly divergent lineage which last shared a most recent common ancestor with the wolf-like
Canis_edwardii
American entrepreneur
dire wolf. Romulus, Remus, and Khaleesi were announced publicly, and he appeared on The Joe Rogan Experience that same month to discuss the dire wolf
Ben_Lamm
2018 asymmetric board game
and was followed by four expansions. A digital version, developed by Dire Wolf Digital, was released in 2020. In Root, 2-4 players compete in an asymmetric
Root_(board_game)
Extinct genus of saber-toothed cat
which S. gracilis preferred soft-skinned browsers. Isotopic studies of dire wolf (Aenocyon dirus) and American lion (Panthera atrox) bones show an overlap
Smilodon
1996 novel by George R. R. Martin
the first chapter—a young boy seeing a beheading and the discovery of dire wolf pups in "summer snows"—and wrote it in a few days. Over that summer, he
A_Game_of_Thrones
1970 album by Grateful Dead
Jerry had been writing; many of them had a decidedly country flavor ('Dire Wolf,' 'Friend of the Devil,' 'High Time,' 'Casey Jones', 'Ripple'), and Jerry
Workingman's_Dead
Name list
the 2018 film Tamizh Padam 2 Khaleesi (wolf) (born 2025), a female genetically modified grey wolf with dire wolf characteristics represented, created by
Khaleesi_(given_name)
Extinct subspecies of the gray wolf
excavations of a large canid, which was comparable in size to North American dire wolf, dating the Late Pleistocene from Aomori and Shizuoka prefectures, however
Japanese_wolf
Extinct species of carnivore
nehringi does not belong to the genus Canis, but may be part of the dire wolf (Aenocyon dirus) lineage, a separate genus, or simply a junior synonym
Canis_nehringi
British actress (born 1996)
Turner's Shocking New Role: Actress Teams Up With Colossal Biosciences For Dire Wolf Revival". The Blast. 2 July 2025. "Engineering the Impossible". SXSW London
Sophie_Turner
2007 American TV series
Smilodon fatalis, dire wolf, short-faced bear, American lion, Bison antiquus, Mexican horse, Columbian mammoth, Megalonyx, gray wolf (cameo), grizzly bear
Prehistoric_Predators
Genus of canines
distinct dire wolf lineage that had convergently evolved a very similar appearance to members of Canis. True members of Canis, namely the gray wolf and coyote
Canis
Calendar year
wolves that are genetically modified to exhibit the characteristics of the dire wolf. April 8 – During a concert performance by Rubby Pérez in Santo Domingo
2025
eldest son of Eddard and Catelyn Stark and the heir to Winterfell. His dire wolf is called Grey Wind. Robb becomes involved in the war against the Lannisters
List of Game of Thrones characters
List_of_Game_of_Thrones_characters
Extinction of large animals at the end of the last Ice Age
Caniformia Canidae Dire wolf (Aenocyon dirus) (C) Pleistocene coyote (Canis latrans orcutti) (C) Megafaunal wolf e.g. Beringian wolf (Canis lupus ssp.)
Late_Pleistocene_extinctions
2025 live album and box set by Grateful Dead
McKernan, Weir, Hunter) – 20:59 Fillmore West – June 7, 1969 – selections: "Dire Wolf" (Garcia, Hunter) – 4:30 "Dupree's Diamond Blues" (Garcia, Hunter) – 4:36
Enjoying_the_Ride
Topics referred to by the same term
Look up dire, diré, or díře in Wiktionary, the free dictionary. Dire may refer to: Diré, a town in Tombouctou Region, Mali Diré Cercle, an administrative
Dire
Family of mammals
cinereoargentus) and the now-extinct dire wolf (Aenocyon dirus). Besides these, there are species endemic to South America: the maned wolf (Chrysocyon brachyurus)
Canidae
American filmmaker
Flare Yes No No Bikini Royale Yes Yes No 2009 Silent Venom Yes No No Dire Wolf Yes No No 2010 Twilight Vamps Yes Yes Executive Bikini Frankenstein Yes
Fred_Olen_Ray
Bulgarian video game artist (1972–2025)
MachineGames' Wolfenstein: The New Order (2014); a visual designer of Dire Wolf Digital's The Elder Scrolls: Legends (2017); and a consultant on Bethesda
Viktor_Antonov_(artist)
Canid native to the southeastern United States
fossil remains of the gray wolf found in the southeastern states. He proposes that following the extinction of the dire wolf, the coyote appears to have
Red_wolf
2016 video game
free-to-play online collectible card video game developed and published by Dire Wolf Digital. Eternal is available for iOS, Android, Xbox One, PCs using Steam
Eternal_(video_game)
American comic book series
Rook: Exodus #7. The arc introduced new Wardens and focused on Rook and Dire Wolf searching for Bloodhound, who may be able to restore Exodus's failing
Rook_(Ghost_Machine)
Subspecies of grey wolf
and fossil North American wolves. The major limb bones of the dire wolf, Beringian wolf, and most modern North American gray wolves can be clearly distinguished
Mexican_wolf
Extinct lineage of the grey wolf
Eastern Asia. Italian wolf, one of the last remaining mDNA haplogroup 2 grey wolves. Subspecies of Canis lupus Pleistocene coyote Dire wolf For a full set of
Pleistocene_wolf
American science fiction drama television series
Castillo, a former runaway whose supposed father, Aaron, died after a dire wolf mauled him. It is later revealed that he kidnapped her when she was a
La_Brea_(TV_series)
Process of producing individual organisms with identical genomes
to Have Brought Back the Dire Wolf". Wired. Retrieved 12 August 2025. Jacobs, Phie (18 April 2025). "What's the deal with dire wolves? Iconic predators
Cloning
Role-playing game system
Cam Banks and initially published by Fandom Tabletop, then acquired by Dire Wolf Digital. Prior versions appeared in the licensed roleplaying games published
Cortex_System
2011 online video game
a 2011 video game based on the Pokémon Trading Card Game developed by Dire Wolf Digital, a studio based in Denver, Colorado. The game was available for
Pokémon_TCG_Online
Novel series by Kathryn Lasky
rogue dire wolves (or outclanners) led by the evil yellow dire wolf Heep. Faolan and Edme also discover that they are reincarnations of two great dire wolves
Guardians_of_Ga'Hoole
2026 Japanese animated fantasy film
hobgoblin servants. Ranga Chikahiro Kobayashi Tyson Rinehart Ranga is a dire wolf serving Rimuru. Treyni Rie Tanaka Jamie Marchi The leader of the dryads
That Time I Got Reincarnated as a Slime the Movie: Tears of the Azure Sea
That_Time_I_Got_Reincarnated_as_a_Slime_the_Movie:_Tears_of_the_Azure_Sea
Extinct genus of canines
(lit. 'dire dog') is an extinct genus of canines that lived between the Pleistocene and early Holocene epochs. The type species is the dire wolf (Aenocyon
Aenocyon
related species such as the dire wolf. 302M, a wolf featured in PBS and National Geographic documentaries 926F (Spitfire), a wolf popular with visitors of
List_of_individual_wolves
1995 video game
the player is made a "21-star general" and transported to the E.S.S. Dire Wolf, where they meet the ship's Flexible Interactive Digital Omnicomputer
Big_Sky_Trooper
Clank!: A Deck-Building Adventure is a 2016 board game published by Dire Wolf Digital, Renegade Game Studios and others. Clank!: A Deck-Building Adventure
Clank!: A Deck-Building Adventure
Clank!:_A_Deck-Building_Adventure
American media franchise based on A Song of Ice and Fire novel series by George R.R. Martin
the first chapter—a young boy seeing a beheading and the discovery of dire wolf pups in "summer snows"—and wrote it in a few days. Over that summer, he
A Song of Ice and Fire (franchise)
A_Song_of_Ice_and_Fire_(franchise)
is a large, silver dire wolf who was born with a splayed paw that has a swirling symbol representing a gyre soul. As a gnaw wolf, he was a member of
List of Guardians of Ga'Hoole characters
List_of_Guardians_of_Ga'Hoole_characters
Lyricist and member of the Grateful Dead (1941–2019)
influence of LSD, and by other hallucinatory experiences: Hunter wrote "Dire Wolf" inspired by a dream after watching a film adaptation of The Hound of
Robert_Hunter_(lyricist)
Robert K.; Larson, Greger; Cooper, Alan; Frantz, Laurent A. F. (2021). "Dire wolves were the last of an ancient New World canid lineage". Nature. 591
Paleobiota of the La Brea Tar Pits
Paleobiota_of_the_La_Brea_Tar_Pits
British actor (born 1959)
Nightmare City 2035 Alex McDowell Cry of the Winged Serpent Griffin 2009 Dire Wolf Sheriff Parker 2013 The Right Regrets Chris Wickham Short film 2015 Those
Maxwell_Caulfield
(2020): Dire Wolf Dune: House Secrets (2021); Portal Games Arrakis: Dawn of the Fremen (2022); Gale Force Nine Dune: Imperium - Uprising (2023): Dire Wolf Dune:
List_of_games_based_on_Dune
2026 American documentary series
ice age. A hibernating short-faced bear (Arctodus) awakens to drive a dire wolf away from her den and cubs. The following spring, mastodons and woolly
Surviving_Earth
Hunting until the prey animal can no longer flee
endurance hunting in extinct species, though potential candidates include the dire wolf Aenocyon dirus due to its similar body shape to modern grey wolves. Non-avian
Persistence_hunting
Extinct subspecies of grey wolf
hybridisation between another grey wolf subspecies and the dire wolf. However, a 2021 study suggested reproductive isolation prevented dire wolves to hybridise with
Canis_lupus_furlongi
American Magic: The Gathering player
Magic: The Gathering Pro Tour player and a game designer for Eternal at Dire Wolf Digital. Chapin has made six Pro Tour top eights, spanning three decades
Patrick_Chapin
Topics referred to by the same term
of North America Coywolf, a wolf-coyote hybrid Dire wolf (Aenocyon dirus), extinct canine of North America Edward's wolf (Canis edwardii), extinct canine
Wolf_(disambiguation)
2024 live album and box set by Grateful Dead
Robert Hunter) – 7:18 "Good Lovin'" (Rudy Clark, Arthur Resnick) – 6:24 "Dire Wolf" (Garcia, Hunter) – 4:38 "Me and My Uncle" > (John Phillips) – 3:08 "Big
Friend of the Devils: April 1978
Friend_of_the_Devils:_April_1978
Subspecies of gray wolf
and fossil North American wolves. The major limb bones of the dire wolf, Beringian wolf, and most modern North American gray wolves can be clearly distinguished
Great_Plains_wolf
Asphalt pit or asphalt lake
The most frequent large mammal found in the La Brea Tar Pits is the dire wolf, one of the most famous prehistoric carnivores that lived during the Pleistocene
Tar_pit
2016 video game
Unlike previous games, there is prehistoric wildlife such as woolly mammoths, dire wolves, cave bears, cave lions, woolly rhinos, brontotherium, Irish elk and
Far_Cry_Primal
1st episode of the 1st season of Game of Thrones
double. In the scene in which the Starks encounter a stag killed by a dire wolf as they return from the execution, an actual animal was used rather than
Winter_Is_Coming
1981 live album by the Grateful Dead
Side A No. Title Writer(s) Recording date and venue Length 1. "Dire Wolf" Jerry Garcia Robert Hunter 10/11/80 Warfield Theatre 3:20 2. "The Race Is On"
Reckoning (Grateful Dead album)
Reckoning_(Grateful_Dead_album)
2011 live album and box set by Grateful Dead
Forest) – 4:23 Disc 2 "Playing in the Band" (Weir, Hart, Hunter) – 11:33 "Dire Wolf" (Garcia, Hunter) – 4:11 Second set: "Good Lovin'" (Clark, Resnick) –
Europe '72: The Complete Recordings
Europe_'72:_The_Complete_Recordings
American musician (born 1954)
Jones" "Cassidy" "China Cat Sunflower" "Cumberland Blues" "Dark Star" "Dire Wolf" "Drums" and "Space" "Eyes of the World" "Fire on the Mountain" "Foolish
Bruce_Hornsby
American actor (1943–2025)
Cabin Massacre! Narrator (voice) Short film 2009 Dire Wolf Col. Hendry Alternative title: Dino Wolf 2012 Blood Fare Professor Meade 2014 Boldly Gone Ben
Gil_Gerard
American musician (1947–2026)
billed as Bobby and Phil. In 2018, Weir formed a band called Wolf Bros. Billed as Bob Weir & Wolf Bros, the group initially was a trio, with Weir on guitar
Bob_Weir
American singer-songwriter and musician
with Lenny Macaluso on an 80s-style rock anthem titled "Happy Birthday Dire Wolf" to celebrate the first birthday of the male wolves created by Colossal
Stan_Bush
2021 zoo-building board game by Mathias Wigge
published Ark Nova: Marine Worlds, an expansion for the base game, and Dire Wolf Digital released Ark Nova Digital as a cross-platform adaptation in 2025
Ark_Nova
Amateur radio data communications protocol
only known complete implementation of v2.2, at this time (2020), is the Dire Wolf software terminal node controller (TNC). AX.25 is commonly used as the
AX.25
Extinct genus of carnivores
ensenadensis Canina (wolf-like canids) see below↓ Canina (wolf-like canids) Canina †Dire wolf (Aenocyon) †Sardinian dhole (Cynotherium) †Trinil dog (Mececyon)
Mesocyon
Species of canine native to North America
exploit the big-game hunting niche left vacant after the extinction of the dire wolf (Aenocyon dirus), as it was rapidly filled by gray wolves, which likely
Coyote
2026 live album by Grateful Dead
Jones" "Cassidy" "China Cat Sunflower" "Cumberland Blues" "Dark Star" "Dire Wolf" "Drums" and "Space" "Eyes of the World" "Fire on the Mountain" "Foolish
Dave's_Picks_Volume_59
American keyboardist, songwriter and singer (1952–1990)
notes, night after night. He was a cross between Gregg Allman and Howlin' Wolf. It was crazy." Mydland's vocals added color to old favorites such as "Cassidy"
Brent_Mydland
PlayStation 4 Xbox One The Elder Scrolls: Legends Windows March 9, 2017 Dire Wolf Digital (2017–2018) Sparkypants Studios (2018–) macOS May 31, 2017 iOS
List of Bethesda Softworks video games
List_of_Bethesda_Softworks_video_games
American science fiction media franchise
Gale Force Nine in 2019. The board game Dune: Imperium was published by Dire Wolf in 2021. In May 2021, a tabletop RPG, Dune: Adventures in the Imperium
Dune_(franchise)
Extinct species of carnivore
greater intelligence. But the abundance of the seemingly intelligent dire wolf at the tar pits calls this hypothesis into question. Despite its rarity
American_lion
2017 free-to-play digital collectible card video game
gain as rewards. The Elder Scrolls: Legends was initially developed by Dire Wolf Digital. The game was revealed at Bethesda's 2015 Electronic Entertainment
The_Elder_Scrolls:_Legends
1987 single by the Grateful Dead
Garcia Robert Hunter Producer John Cutler Grateful Dead singles chronology "Dire Wolf" (1981) "Touch of Grey" (1987) "When Push Comes to Shove" (1987)
Touch_of_Grey
1996 live album by Grateful Dead
(electric): "China Cat Sunflower" → "I Know You Rider", "Me and My Uncle", "Dire Wolf", "Smokestack Lightning"[b] Second set (acoustic): "The Monkey and the
Dick's_Picks_Volume_4
Season of television series
Warden of the North, executes him for desertion. Later, six orphaned dire wolf pups are found and one given to each Stark sibling, including Ned's bastard
Game_of_Thrones_season_1
2022 studio album by Molly Tuttle
released on December 2, 2022, featured four bonus tracks - a cover of "Dire Wolf" by Grateful Dead, a rendition of the traditional folk song "Cold Rain
Crooked_Tree_(album)
Paleontological research site in Los Angeles
Retrieved May 9, 2021. Ojibwa (December 13, 2017). "Paleontology 101: The Dire Wolf". Daily Kos. Kos Media, LLC. Retrieved May 9, 2021. Udurawane, Vasika;
La_Brea_Tar_Pits
Dice-drafting board game
Facades were released between 2019 and 2022. A digital version developed by Dire Wolf Digital was published as an app in 2020. In 2023, a legacy edition of
Sagrada_(board_game)
Extinct Ice-Age European subspecies of wolf
domestic dog C. l. familiaris descended from these extinct lineages. Dire wolf Kempe, Stephan; Döppes, Doris (2009). "Cave Bear, Cave Lion and Cave Hyena
Cave_wolf
National park in New Mexico, United States
lakebed that appeared to be footprints. During the 2010s, footprints of a dire wolf were discovered beside ancient seeds dated to more than 18,000 years ago
White_Sands_National_Park
American guitarist and singer (1942–1995)
(1976-1977). On September 28, 1977, Irwin delivered the refurbished Wolf back to Garcia. The wolf sticker which gave the guitar its name had now been inlaid into
Jerry_Garcia
American fantasy drama TV series (2011–2019)
meeting 'de-extincted' dire wolves". The Independent. Retrieved April 30, 2025. Pener, Degen (April 22, 2025). "The Dire Wolf's Hollywood DNA". The Hollywood
Game_of_Thrones
2016 video game
inventory UI audio and block sound pickup/place, blood splatter on bear/dire wolf, prefab editor improvements, Twitch menu enhancements, AI investigation/range
7_Days_to_Die
Process which formed the domestic dog
increase in tooth breakage that is similar to that seen in the extinct dire wolf. This suggests that these either often processed carcasses, or that they
Domestication_of_the_dog
North American ice age, 9,000–16,000 BCE
November 18, 2016. United States National Park Service: Dire Wolf https://www.nps.gov/articles/000/dire-wolf.htm "Gray Wolves in Washington State: Questions and
Vashon_Glaciation
January 24, 2025 The Elder Scrolls: Legends WIN, OSX, iOS, DROID DCCG Dire Wolf Digital, Sparkypants Studios Bethesda Softworks January 30, 2025 EA Sports
2025_in_video_games
American Magic: The Gathering player
but ended his tenure as editor in 2012 to work as a game designer at Dire Wolf Digital, specifically on Eternal, and in 2021 LSV joined Good Luck Games
Luis_Scott-Vargas
American geneticist (born 1954)
live birth of two "dire wolves" where gray wolf cells received 20 unique gene edits, including 15 edits that directly matched dire wolf DNA and notably an
George_Church_(geneticist)
Paleozoographic event resulting from the formation of the Isthmus of Panama
Bibcode:2005JSAES..20..105S. doi:10.1016/j.jsames.2005.07.005. "Canis dirus (dire wolf)". Paleobiology Database. Archived from the original on 2021-02-15. Retrieved
Great_American_Interchange
American writer and television producer (born 1948)
Writers". Snider, Mike (April 8, 2025). "George R.R. Martin got to hold a dire wolf. Extinct animal's 'rebirth...has stirred me'". USA Today. "George R.R
George_R._R._Martin
Tabletop games based on A Song of Ice and Fire franchise
2-6 Board game Hasbro Game of Thrones: Oathbreaker 2019 5-8 Board game Dire Wolf Funkoverse Strategy Game: Game of Thrones 100 2020 2-4 Board game Funko
List of A Song of Ice and Fire tabletop games
List_of_A_Song_of_Ice_and_Fire_tabletop_games
Fossil site in San Diego County, California
phalanx bones, 2 sesamoids and over 300 other bone fragments. Remains of dire wolf, horse, camel, mammoth and ground sloth were also found. Five cobbles
Cerutti_Mastodon_site
Fauna of the US state of Florida
or extirpated; such as the Florida short-faced bear, Florida black wolf, Dire wolf, Dexteria floridana, Florida bog lemming, Long-nosed peccary, Caribbean
Fauna_of_Florida
2026 live album by Grateful Dead
Jones" "Cassidy" "China Cat Sunflower" "Cumberland Blues" "Dark Star" "Dire Wolf" "Drums" and "Space" "Eyes of the World" "Fire on the Mountain" "Foolish
Dave's_Picks_Volume_58
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